Edinburgh Zoo has confirmed its resident giant panda will not give birth to a cub this year.

Zookeepers say Tian Tian's hormone levels and general behaviour signal the end of the animal's breeding cycle for the rest of the year.

The two pandas arrived at the zoo almost five years ago and were celebrated as the first giant pandas to live in the UK for 17 years.

Tian Tian, which means Sweetie, and male Yang Guang, meaning sunshine, have so far failed to produce a cub since their arrival to Edinburgh Zoo.

Iain Valentine, director of giant pandas at the Royal Zoological Society of Scotland (RZSS), said: "It now appears clear that Tian Tian, our resident female giant panda, will not give birth to cubs this year.

"Tian Tian's hormone levels are returning to normal and her behaviours signal the end of her breeding cycle for this year.

"Over the next few weeks we will conduct a thorough review of the scientific data and our processes to ensure we learn from this year's breeding season."